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How was it concluded that genes are loca...

How was it concluded that genes are located on chromosomes ?

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Morgan confirmed Mendelian laws of inheritance and the hypothesis that genes are located on chromosomes. Morgan had discovered that eye colour in Drosophila expressed a sex-linked trait. All first-generation offspring of a mutant whiteeyed male and a normal red-eyed female would have red eyes because every chromosome pair would contain at least one copy of the X chromosome with the dominant trait.
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