Shrubs and trees originating from temperate regions typically exhibit discernible growth layers commonly known as growth rings or annual rings within their secondary xylem. These annual rings, visible as distinctive concentric circles in the cross-section of a tree trunk, serve as a fundamental aspect of understanding the life history and growth patterns of individual trees. The formation of these rings is intricately linked to the dynamic activity of the vascular cambium, a layer of meristematic tissue responsible for the secondary growth in woody plants. By studying these annual rings, scientists gain insights into the cyclical yearly growth cycles of trees.
Each annual ring comprises two distinct growth rings—one of autumn wood and one of spring wood. These rings collectively represent one year of the tree's growth. Counting the number of rings from the base of the stem provides a reliable estimate of the tree's age. Increment borer instruments facilitate the extraction of core samples without causing significant harm, allowing for accurate age determination.
The study of annual rings, known as dendrochronology, extends beyond age determination. It offers valuable insights into past climates, growth patterns, and environmental shifts. Deciduous trees, which shed their leaves annually, typically present clearer annual rings compared to evergreen counterparts.
In certain conditions, such as drought stress during a growing season, trees may form more than one ring, termed pseudo annual rings. Overall, annual rings serve as a tangible historical record embedded within the structure of trees, providing a wealth of information about their dynamic relationship with the environment.
The formation of annual rings in trees is governed by the dynamic activity of the cambium, a layer of meristematic tissue that undergoes fluctuations throughout the year. These annual rings are characterized by distinct growth layers, with two primary types: spring wood (or early wood) and autumn wood (also referred to as late wood).
Spring wood, also known as early wood, is a component of annual rings formed during periods of active cambium activity in spring or summer. This wood is characterized by larger cells, thinner cell walls, and wider lumens. It is generally lighter in color and exhibits lower density compared to autumn wood. The formation of spring wood contributes to the outward expansion of the annual ring during periods of increased growth.
Autumn wood, or late wood, is another component of annual rings formed when cambium activity is reduced, typically in winter or autumn. Cells formed during this period are smaller, have thicker walls, and narrower lumens. Autumn wood is darker in color and has a higher density due to the compact arrangement of cells. The combination of spring wood and autumn wood forms the complete annual ring, representing one year of the tree's growth.
(Session 2025 - 26)