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How was it concluded that genes are loca...

How was it concluded that genes are located on chromosomes?

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Chromosome theory of inheritance was proposed by Sutton and Boveri independently in 1902. The theory believes that chromosomes are vehicles of heriditary information, prossess Mendelian factors of ganes and it is the chromosomes which segregate and assort independently during transmission from one generation to the next.
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NCERT EXEMPLAR-PRINCIPLE OF INHERITANCE AND VARIATIONS -Principle Of Inheritance And Variations
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