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The diagram shows a bivalent at prophase...

The diagram shows a bivalent at prophase -I of meiosis. Which of the four chromatids can cross over ?

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The homologous chromosomes lie parallel to each other in leptotene stage. Each chromosome has four chromatids and are bivalent. The non-sister chromatids of homologous chromosomes cross over in pachytene stage of prophase-I
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