Anther wall consists of
(a) One cell thick outer protective layer, epidermis. The thinWalled epidermal cells between two pollen sacs constitute stomium, which is useful for the dehiscence of pollen sacs.
(b) Endothecium, a layer of radially expanded cells present below the epidermis. These cells have hygroscopic-fibrous. thickenings of cellulose and help in the dehiscence of pollen sac
(c) Middle layers present below the endothecium consists of 1-5 layers of thin walled cells.
(d) Tapetum is the innermost nutritive layer of anther wall, which encircles the sporogenous tissue. The cells in this layer are large ,thin walled , bi - or multinucleate and nourish the developing sporogenous tissue.